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Devadatta is a lion

A famous example of secondary meaning involving properties (gauṇavṛtti) is "Devadatta is a lion," explained by Kumārila Bhaṭṭa in his TV to mean that Devadatta has properties of a lion, such as courage.

However, in the PāliJavasakuṇa Jātaka, we hear the Buddha quite literally identify Devadatta as being a lion, in a previous birth, who showed ingratitude to the Buddha who was at that time born as a woodpecker. (This line doesn't seem to appear in the Sanskrit Jātakamāla.)

Could it be the case that the comparison between Devadatta and a lion in the brahminical context is connected to this story? I wasn't able to find anyone suggesting this in a quick search, though there are independent treatments of the brahminical metaphor (connoting courage) and the Buddhist story (exemplifying greed or ingratitude). I don't know when it emerged in the brahminical context, but since Devadatta is antagonistic towards the Buddha, I could see Kumārila taking the identification an…

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